Form: S-8

Securities to be offered to employees in employee benefit plans

August 13, 2025

S-8 S-8 EX-FILING FEES 0001200375 CODEXIS, INC. N/A Fees to be Paid 0001200375 2025-08-11 2025-08-11 0001200375 1 2025-08-11 2025-08-11 iso4217:USD xbrli:pure xbrli:shares

Calculation of Filing Fee Tables

S-8

CODEXIS, INC.

Table 1: Newly Registered Securities

Security Type

Security Class Title

Fee Calculation Rule

Amount Registered

Proposed Maximum Offering Price Per Unit

Maximum Aggregate Offering Price

Fee Rate

Amount of Registration Fee

1 Equity Common Stock, par value $0.0001 per share Other 8,000,000 $ 2.78 $ 22,240,000.00 0.0001531 $ 3,404.94

Total Offering Amounts:

$ 22,240,000.00

$ 3,404.94

Total Fee Offsets:

$ 0.00

Net Fee Due:

$ 3,404.94

Offering Note

1

(1) Pursuant to Rule 416(a) under the Securities Act of 1933, as amended (the "Securities Act"), this Registration Statement shall also cover any additional shares of common stock of Codexis, Inc. (the "Company") that become issuable under the Codexis, Inc. 2019 Incentive Award Plan, as amended, by reason of any stock dividend, stock split, recapitalization or other similar transaction effected that results in an increase in the number of outstanding shares of the common stock of the Company. (2) The offering price per unit is estimated in accordance with Rules 457(c) and (h) under the Securities Act solely for the purpose of calculating the amount of the registration fee, based on the average of the high and low sale prices of the common stock reported on the NASDAQ Global Select Market on August 8, 2025.

Table 2: Fee Offset Claims and Sources ☑Not Applicable
Registrant or Filer Name Form or Filing Type File Number Initial Filing Date Filing Date Fee Offset Claimed Security Type Associated with Fee Offset Claimed Security Title Associated with Fee Offset Claimed Unsold Securities Associated with Fee Offset Claimed Unsold Aggregate Offering Amount Associated with Fee Offset Claimed Fee Paid with Fee Offset Source
Rule 457(p)
Fee Offset Claims N/A N/A N/A N/A N/A N/A N/A N/A N/A N/A N/A N/A
Fee Offset Sources N/A N/A N/A N/A N/A N/A N/A N/A N/A N/A N/A N/A